Statistics, mathematical modelling, and epidemiology with a fair bit of ecology and immunology thrown in
| 1998-2001 | Fellow of King's College Cambridge and NERC Advanced Research Fellow in Zoology |
| 1996-1998 | NERC postdoctoral researcher, Plant Sciences and Zoology, Cambridge. |
| 1990-1996 | Wellcome postdoctoral researcher, Biology, Imperial College and Zoology, Oxford |
| 1987-1990 | PhD in Applied Mathematics, Imperial College London |
| 1986-1987 | Certificate of Advanced Study in Mathematics, Cambridge. |
| 1983-1986 | BA in Mathematics, King's College Cambridge. |